The other species to which I have to direct attention is the Kallima paralekta, a butterfly of the same family group as our Purple Emperor, and of about the same size or larger. Its upper surface is of a rich purple, variously tinged with ash colour, and across the forewings there is a broad bar of deep orange, so that when on the wing it is very conspicuous. This species was not uncommon in dry woods and thickets, and I often endeavoured to capture it without success, for after flying a short distance it would enter a bush among dry or dead leaves, and however carefully I crept up to the spot I could never discover it until it would suddenly start out again and then disappear in a similar place. If at length I was fortunate enough to see the exact spot where the butterfly settled, and though I lost sight of it for some time, I would discover that it was close before my eyes, but that in its position of repose it so closely resembled a dead leaf attached to a twig as almost certainly to deceive the eye even when gazing full upon it. I captured several specimens on the wing, and was able fully to understand the way in which this wonderful resemblance is produced.
The end of the upper wings terminates in a fine point, just as the leaves of many tropical shrubs and trees are pointed, while the lower wings are somewhat more obtuse, and are lengthened out into a short thick tail. Between these two points there runs a dark curved line exactly representing the midrib of a leaf, and from this radiate on each side a few oblique marks which well imitate the lateral veins. These marks are more clearly seen on the outer portion of the base of the wings, and on the innerside towards the middle and apex, and they are produced by striae and markings which are very common in allied species, but which are here modified and strengthened so as to imitate more exactly the venation of a leaf. The tint of the undersurface varies much, but it is always some ashy brown or reddish colour, which matches with those of dead leaves. The habit of the species is always to rest on a twig and among dead or dry leaves, and in this position with the wings closely pressed together, their outline is exactly that of a moderately-sized leaf, slightly curved or shrivelled.
The tail of the hind wings forms a perfect stalk, and touches the stick while the insect is supported by the middle pair of legs, which are not noticed among the twigs and fibres that surround it. The head and antennae are drawn back between the wings so as to be quite concealed, and there is a little notch hollowed out at the very base of the wings, which allows the head to be retracted sufficiently. All these varied details combine to produce a disguise that is so complete and marvellous as to astonish everyone who observes it; and the habits of the insects are such as to utilize all these peculiarities, and render them available in such a manner as to remove all doubt of the purpose of this singular case of mimicry, which is undoubtedly a protection to the insect.
Its strong and swift flight is sufficient to save it from its enemies when on the wing, but if it were equally conspicuous when at rest it could not long escape extinction, owing to the attacks of the insectivorous birds and reptiles that abound in the tropical forests.
A very closely allied species, Kallima inachis, inhabits India, where it is very common, and specimens are sent in every collection from the Himalayas. On examining a number of these, it will be seen that no two are alike, but all the variations correspond to those of dead leaves. Every tint of yellow, ash, brown, and red is found here, and in many specimens there occur patches and spots formed of small black dots, so closely resembling the way in which minute fungi grow on leaves that it is almost impossible at first not to believe that fungi have gown on the butterflies themselves!
If such an extraordinary adaptation as this stood alone, it would be very difficult to offer any explanation of it; but although it is perhaps the most perfect case of protective imitation known, there are hundreds of similar resemblances in nature, and from these it is possible to deduce a general theory of the manner in which they have been slowly brought about. The principle of variation and that of "natural selection," or survival of the fittest, as elaborated by Mr. Darwin in his celebrated "Origin of Species," offers the foundation for such a theory; and I have myself endeavoured to apply it to all the chief cases of imitation in an article published in the "Westminster Review" for 1867, entitled, "Mimicry, and other Protective Resemblances Among Animals," to which any reader is referred who wishes to know more about this subject.
In Sumatra, monkeys are very abundant, and at Lobo Kaman they used to frequent the trees which overhang the guard-house, and give me a fine opportunity of observing their gambols. Two species of Semnopithecus were most plentiful--monkeys of a slender form, with very long tails. Not being much shot at they are rather bold, and remain quite unconcerned when natives alone are present; but when I came out to look at them, they would stare for a minute or two and then make off. They take tremendous leaps from the branches of one tree to those at another a little lower, and it is very amusing when a one strong leader takes a bold jump, to see the others following with more or less trepidation; and it often happens that one or two of the last seem quite unable to make up their minds to leap until the rest are disappearing, when, as if in desperation at being left alone, they throw themselves frantically into the air, and often go crashing through the slender branches and fall to the ground.